Marc and Betty compete against each other for the Young Editors Training Program but only one spot is available. Daniel learns he is drawn towards Molly. Wilhelmina attempts to get Connor to notice her.
The sudden death of a Mode editor leaves Betty and Marc competing for the same job. Meanwhile, Wilhelmina demands that Claire tender her resignation at a big awards ceremony … or she’ll reveal Claire’s dark secret to Cal Hartley; Matt lets his jealousy over Henry get the best of him; and Justin nervously waits to see if he got into performing arts school.
Just as Betty agrees to move in with Matt, Henry (guest star Christopher Gorham) pays a visit to NYC, forcing Betty to admit she still has feelings for her former fiance. Meanwhile, Betty and Marc wait to see if their YETI editor interviews resulted in actual jobs; Daniel’s transformation from playboy to devoted husband makes him a media darling; and Wilhelmina cozies up to Victoria Hartley to find out just what kind of connection Claire Meade has to Cal Hartley.
Betty teams with Matt and Marc for their all-important final YETI project, a photo shoot with Grammy winner Adele, but Matt’s slacking off could jeopardize everything they’ve worked for. Meanwhile, love is everywhere as two wedding proposals are made, but only one couple will walk down the aisle; Claire and Wilhelmina square off over their roles at Mode; and Matt’s mom (guest star Christine Baranski) asks Betty for a favor.
Just as it looks like billionaire Calvin Hartley (guest star David Rasche) is going to give Mode the bailout it needs so the magazine won’t go under, baby William is kidnapped - and Betty is dragged into the ensuing drama. Meanwhile Archie (guest star Ralph Macchio) asks Hilda to be part of his latest political campaign.
When Mode gets reposessed, Betty and Daniel decide to take drastic actions. Meanwhile, Christina is stunned when she learns that the Meade/Slater baby may be her biological son and not Wilhelmina’s, and Matt makes a life-changing proposition to Betty when the Suarez family meets Matt’s father at his renowned Easter egg hunt!
Daniel and Wilhelmina make Modes issue based on sex to spice things up. Matt is hesitant to take the next step with Betty. Marc and Amanda provide Betty with a makeover.
Betty comes face to face with Matt’s mother. Wilhelmina devizes a way to make money fast whilst Daniel deals with Molly. The Suarez family attempt to set Hilda and Archie up on a date.
The Suarez family attempt to buy their rented house from their landlord. Ignacio goes head to head with Frankie Burrata, a celebrity chef. While everyone in Mode comes to deal with their shock.
A YETI assignment grants a shocking look at Mode’s financial history for Marc and Betty, who also copes with news that Henry has posted on the Internet and deals with increased attention from a previous YETI partner, Matt (Daniel Eric Gold).
Fashion Week in New York City makes Betty’s life more hectic than usual as she helps an eccentric designer put on a show and partners up for a YETI assignment with someone who works at a sports magazine. Meanwhile, Wilhelmina worries about her new relationship’s effect on her career, and Daniel hopes to see another side of Molly.
It’s Claire’s 60th birthday and Betty makes her a video which reveals some unintentional footage of Mrs Meade. Also, Daniel and Wilhelmina are finding it hard having someone new in their lives.
Betty comes up with a solution to deal with her personal problems but she has her sexy neighbour Jesse who keeps her distracted from her goals. Daniel does his best to cover his tracks as Connor is adament on unveiling Molly’s admirer.
Betty finds herself stuck between work and home life as she tries her best to appease both Hilda; Daniel, Molly, Connor and Wilhelmina.
Betty fights an uphill battle to make a good impression on a YETI editor (Bernadette Peters), and meets her counterpart (Nikki Blonsky) at Elle magazine as the publication’s rivalry with Mode continues. Elle editors Robbie Myers and Joe Zee appear as themselves.
Amanda and Betty are forced to work together for an article called “How I Blew Ten Grand Without Spending a Dime.”